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HIERARCHICAL CLASSIFICATION OF THE PHYLUM BRACHIOPODA

HIERARCHICAL CLASSIFICATION OF THE PHYLUM BRACHIOPODA (DUMERIL 1806)

EUKARYA> UNIKONTA> OPISTHOKONTA> ANIMALIA> METAZOA> BILATERIA> PROTOSTOMATA> SPIRALIA> TROCHOZOA> BRACHIOZOA> BRACHIOPODA

SUBPHYLUM LINGUILIFORMEA

Shell valves, normally of calcium phosphate and chitin, held together solely by muscles; lophophore without a supporting skeleton; gut with an anus; peduncle with muscles and coelomic cavity; body cavity forms schizocoelically (text with tooltip) Schizocoelic (adj.) describes a type of true coelom that develops by the separation of mesoderm tissue. from mesoderm produced enterocoelically.

CLASS LINGULATA (3 ORDERS)

  • Dorsal and ventral valves separated; marginal setae continuous; statocysts in larvae and adults.
    • Early Cambrian – Recent
    • Discinida, Discinisca, Discradisca, Pelagodiscus.
    • Glottida, Lingula
  • Margins without setae.
    • Early Cambrian – Late Orodovician

SUBPHYLUM CRANIIFORMEA

Valves mineralized with calcite; no peduncle; ventral valve cemented to hard substrate.

CLASS CRANIATA (3 ORDERS)

  • Early Cambrian – Recent
    • Crania, Craniscus, Neoancistrocrania, Valdiviathyris, Neocrania.

SUBPHYLUM RHYNCHONELLIFORMEA

Shell valves of calcium carbonate, with hinges of teeth and sockets; lophophore usually with supporting skeleton; gut without anus; peduncle without coelomic cavity or muscles; body cavity enterocoelic; peduncle, if present, emerging through slit or notch in ventral shell.

  • Shells inarticulated.
    • Early Cambrian – Permian
  • Early Cambrian – Middle Cambrian
  • Early Cambrian – Middle Cambrian
  • Middle Cambrian – Late Permian

CLASS RHYNCHONELLATA (9 ORDERS)

  • Distinctive articulation structures.
    • Early Cambrian – Recent
    • Aulites, Cryptopora, Hemithiris, Pemphixina, Notosaria
    • Compsothyris, Frieleia, Grammetaria, Sphenarina, Abyssorhynchia, Hispanirhynchia, Manithyris, Parasphenarina, Neorhynchia, Tethyrhynchia
    • Acanthobasiliola, Striarina, Basliola, Basilioella, Rhytirhynchia
    • Bouchardia, Fallax, Septicollarina, Laurinia, Ecnomiosa, Kraussina, Megerlia, Megerlina, Pumilus, Frenulina, Jolonica, Pictothrys, Shimodaia, Argyrotheca, Joania, Megathris, Leptothyrella, Phaneropora, Amphithyris, Annuloplatidia, Platidia, Holobrachia, Simplicithrys
    • Cancellothrys, Murravia, Surugathyris, Terebratulina, Agulhasia, Bathynanus, Eucalathis, Melvicalathis, Nanacanathis, Notozyga, Abyssothyris, Acrobelesia, Xenobrochus, Dyscolia, Goniobrochus, Dallithrys, Kanakythryris, Stenosarina, Gryphus, Acrobrochus, Liothyrella, Arctosia, Dolichozygus, Dysedrosia, Erymnia, Tichosina, Zygonaria, Lacazella, Osperyella, Pajaudina, Kakanuiella, Minutella, Thecidellina
LITERATURE CITED

Barnes, R. D. 1980. Invertebrate Zoology. Saunders College/Holt, Rinehart and Wilson, Philadelphia.

Barnes. R. S. K. 1984a. Kingdom Animalia. IN: R. S. K. Barnes, ed. A Synoptic Classification of Living Organisms. Sinauer Associates, Inc., Sunderland, MA. pp. 129-257.

Brusca, R. C. and G. J. Brusca. 2003. Invertebrates. Sinauer Associates, Inc. Sunderland, Mass.

Buchsbaum, R. 1938. Animals Without Backbones, An Introduction to the Invertebrates. The University of Chicago Press. Chicago.

Hickman, C. P. 1973. Biology of the Invertebrates. The C. V. Mosby Company. Saint Louis.

Margulis, L. and K. Schwartz. 1998. Five kingdoms, an illustrated guide to the phyla of life on earth. 3rd Edition. W. H. Freeman and Company. New York.

Meglitsch, P. A. and F. R. Schramm. 1991. Invertebrate Zoology. Oxford University Press, New York, Oxford.

Nielsen, C. 2001. Animal Evolution: Interrelationships of the Living Phyla. 2nd Edition. Oxford University Press. Oxford.

Pechenik, J. A. 2005. Biology of the Invertebrates. McGraw-Hill. New York.

Ruppert, E. E. and R. D. Barnes. 1994. Invertebrate Zoology. 6th edition. Saunders. Ft Worth, TX.

Ruppert, E. E., R. S. Fox, and R. D. Barnes. 2004. Invertebrate Zoology: A Functional Evolutionary Approach. Seventh Edition. Thomson, Brooks/Cole. New York. pp. 1-963.

Storer, T. I. and R. L. Usinger. 1965. General Zoology. 4th Edition. McGraw-Hill Book Company. New York.

Tudge, C. 2000. The Variety of Life, A Survey and a Celebration of all the Creatures That Have Ever Lived. Oxford University Press. New York.
By Jack R. Holt. Last revised: 01/28/2012
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